It could be the 4th engine or it could be the 20th engine. These are possibilities, but what's at issue is the probability. Based on this particular car's performance and referencing it against the Motor Trend car (which was independently dyno'ed), I think it's safe to say this car most certainly did not have 100 more hp than the Motor Trend car, which most people accept as performing on a level expected with the claimed hp specification (if anything, the consensus has been that it's a bit slow).
If this was a worn out test rig, then we'd have seen a lot more press of it. A red Lexus LFA seen lapping the 'Ring or on the autobahn long enough to go through 4 engines would be pretty easy to spot. Lexus's image is riding on this car and I think they know it would be pretty foolish to deliver a product that were signficantly slower.
For the record, the red LFA was first seen briefly as far back as March 29th, in Japan. It was later put on static display at a car show in Japan. It wasn't spotted again until the Goodwood session and to my knowledge, no red LFA has since been seen back in Japan. I'm also fairly certain the whereabouts of the other LFA's can still be accounted for.
As to the possibility of the tripmeter, it's possible. But not likely. From the Motor Trend dyno video, we can see there is definitely a tripmeter labeled, when the function is enabled.